Zeke Yeager, a divisive character in Attack on Titan, betrays his parents and friends, leading to devastating consequences. Zeke's ruthless actions, including killing beloved characters and ...

Zeke's predecessor as the holder of the Beast Titan was his mentor, Tom Ksaver, who was an Eldian researcher, member of Marley's Warrior Unit, and a member of Marley's Titan Biology Research Society.

Characters like Vegeta and Light Yagami became anti-heroes with morally gray motivations. Recent anime introduced deeply human and sympathetic villains like Shigaraki Tomura and Zeke Yeager.